- Vakinankaratra Region, Ambatolampy Dist., Manjakatompo forest station2
- Vanrhynsdorp. Klawer2
- Vohipaho Reserve2
- (Northe of) Birnin Gwari1
- (South-East). Zombitse National Park, park entrance road near RN71
- ANOSY-Amboasary. Flanc de colline, dans la forêt sèche à 31 km au nor-est d'Amboasary.1
- ANOSY-Amboasary. Forêt sèche é31 km au nord-est d'Amboasary, sur le bord de la route traversant la forêt.1
- Africa, Central1
- Alaotra Mangoro, Antampimbato, Analamay. Forest without cuirasse. Collected with F. Edmond, B.A. Ratodimanana, S. Randrianasolo & J.F. Randriamitantsoa.1
- Alaotra Mangoro, Moramanga, Ambohibary, Ampitambe, Ambatovy. Andranovery edge of forest. Collected with S. Randrianasolo, R. Bernard, B.A. Ratodimanana & F. Edmond.1
- Alaotra Mangoro, Moramanga, Andasibe, Menalamba, Antsahalava. Antsahalava forest. Collected with H.R. Radona, B.A. Ratodimanana, J.F. Randriamitantsoa & A. Rakotondrafara.1
- Albertville1
- Algeria1
- Along Iringa-Kitonga road ~25 km from Iringa.1
- Along RIP 118, within Andohahela National Park, just past Manangotry Pass.1
- Along RN 13 between Amboasary Sud and Ranopiso. East of large sisal plantation1
- Along RN 7.1
- Along Route Nationale 6, at the top of a local high point1
- Along road from Fort Dauphin towards Petriky.1
- Along road from RN 13 to Tsimelahy, leading into Parcel II of Andohahela National Park.1