- Analamanga3
- Village of Ambovomamy; Route Nationale 6 at PK 178; south of the Sofia River, on northwest side of highway across from the village3
- Along road from RN 13 to Tsimelahy, leading into Parcel Ⅱof Andohahela National Park2
- Ankarana National Park; Circuit Benavony.2
- Boeny2
- Lourenço Marques2
- Mantadia National Park, along road going North into the park.2
- Maputo2
- Mozambique2
- On road going to the NE past the southern end of the runway at the Majunga Airport along road to Antsanitia.2
- Past the entrance to Parc National Andohahela. Tsimelahy road.2
- RN 6, just north of the turnoff to Belinta.2
- Ranomafana National Park, lower part of Talatakely trail system2
- Road from Route Nationale 2 to Lakato.2
- Vakinankaratra Region, Ambatolampy Dist., Manjakatompo forest station2
- Vohipaho Reserve2
- Alaotra Mangoro, Antampimbato, Analamay. Forest without cuirasse. Collected with F. Edmond, B.A. Ratodimanana, S. Randrianasolo & J.F. Randriamitantsoa.1
- Alaotra Mangoro, Moramanga, Ambohibary, Ampitambe, Ambatovy. Andranovery edge of forest. Collected with S. Randrianasolo, R. Bernard, B.A. Ratodimanana & F. Edmond.1
- Alaotra Mangoro, Moramanga, Andasibe, Menalamba, Antsahalava. Antsahalava forest. Collected with H.R. Radona, B.A. Ratodimanana, J.F. Randriamitantsoa & A. Rakotondrafara.1
- Along RIP 118, within Andohahela National Park, just past Manangotry Pass.1