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Place
- Firondronana de Mananara-Nord, Ivontaka-Sud; forêt à 100 m de la mer; parcelle 4.1
- Mantadia National Park, along trail starting from the northernmost parking area of road going North into park.1
- Masoala National Park. Ridge NNE of summit of Ambohitsitondroinan'Mahalevona, SSE of village of Mahalevona. Low perhumid forest along trail.1
- Moramanga, Analamazaotra1
- Moramanga, Andasibe, Analamazaotra-Périnet RS, Circuit Indri 1.1
- Moramanga, Andasibe, Berano. Ambatovy, Ampangadiantrandraka forest.1
- Moramanga, Andasibe, Parc National Analamazaotra-Périnet, Andranomadio.1
- Moramanga, Andasibe, Parc National Mantadia, Falierana, Sahandambo.1
- Moramanga, Andasibe, Parc National d'Analamazaotra-Périnet.1
- Moramanga, fokontany Ampangalatsara. Ambodimandresy forest. Preserved in alcohol.1
- Road from Brickaville to Ambila Lemaitso.1
- Road to Analalava, west of Foulpointe.1
- Région Alaotra-Mangoro, District de Moramanga, Commune Andasibe, FKT Ampangalatsara, forêt d'Ambodimandresy, Hotel Restaurant Eulophiella.1
- Soanierana Ivongo1
- Station forestière de Tampolo, 10 km north of Fenoarivo, east of RN 5. Plant preserved in alcohol, collected under MBG second Field Botanist Training program with students from ESSA; part of "Inventaire Biologique Tampolo 1997."1
- Torotorofotsy forest, Fokotany Ampitambe, Ambatovy, commune rurale Ambohibary, district Moramanga1
- Vicinity of Hotel Plage Dauphin, Ambila-Lemaitso.1
- Vohibinany (Brickaville), Ampitabe, 8 km from Antsampanana towards Vatomandry, Analapetraka forest1