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- Ankarana National Park; Circuit Benavony.2
- RN 6, just north of the turnoff to Belinta.2
- Reserve Naturelle Marojejy, along the trail to the summit of Marojejy Est, below the third camp2
- Ampasindava, forêt de Bongomihiravavy.1
- Andratamarina, Sambava.1
- Ankarana National Park; along the Maeva trail, on tsingy, on bare rock with only a little accumulation of soil. At the top of the hill where the trail levels off.1
- Antsaha, Mantazanina, Ambalavy, Antanandavahely, Ambohitralanana, Antalaha1
- Betsiaka. Andavakoera forest. Approximately ½ way up the slope to the top of the hills.1
- Diego, Diana, Ambilobe, Manambato, Antsavahalany. Forêt dense humide.1
- Fivondronana d'Andapa. Réserve spécial d'Anjanaharibe Sud. Aux environs du sommet. Plantes préservées dans l'alcool.1
- Marojejy RNI, au nord d'Andapa. Andrahanjo, Sambava. Forêt dense humide, sol sableux. Echantillon préservé en alcool.1
- Montagne d'Ambre, partie centrale. A 390 m du point cote 1445, au 179?1
- Montagne d'Ambre, versants ouest.1
- RN 6 east of Ampampamena and south of Ambatomainty.1
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