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- Guatemala12
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- Gualan8
- Along a path about 3/4 mile from the river on opposite side from Gualan1
- Contact UM Herbarium1
- Gualan; Climbing on bushes 10 ft. high along fence on west side of the Motagua1
- along the Gualan River1
- along upper reaches of Río Sitio Nuevo, between Santa Rosalía and first waterfall1
- ravine bordering Quebrada Alejandria, summit of Sierra de Las Minas Vicinity of Finca Alejandria1