Place
- Central Paraguay.1
- Cerro León (Lagerenza), oeste del cerro, Chaco1
- Cnia. Pindo, camino entre Itaquyry y Curuguati1
- Cordillera de Altos1
- Cordillera de Ybytyruzú. Valley between Cerro Peró Acati and Cerro Muy Muy1
- Cordillerae Centralis. Cerros de Tobaty region.1
- Cordillere de Mbatobi, near Paraguari1
- Corrientes River Region. Sierra de Maracayu1
- Estancia Zalazar. Laguna Zalazar1
- General Bernardio Caballero1
- Guairá1
- In Dumets Campestribus Pr. Bellavista (Apa)1
- In Forest near Tobaty1
- In Viciniis Caaguazú1
- In altaplanitie et declivibus Sierra de Amambay1
- In colle Tobaty1
- Isla Yacyreta. Collecting area ca. 100 square m1
- Mbaracayu Biosphere reserve. aguara nu. collecting area around two hectares1
- Mbaracayú Biosphere Reserve. Aguará ñu.1
- Mbaracayú Biosphere Reserve. Aguará-Ñu1