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- Ankarana National Park; Circuit Benavony.2
- RN 6, just north of the turnoff to Belinta.2
- Ampasindava, forêt de Bongomihiravavy.1
- Ankarana National Park; along the Maeva trail, on tsingy, on bare rock with only a little accumulation of soil. At the top of the hill where the trail levels off.1
- Betsiaka. Andavakoera forest. Approximately ½ way up the slope to the top of the hills.1
- Montagne d'Ambre, partie centrale. A 390 m du point cote 1445, au 179?1
- Montagne d'Ambre, versants ouest.1
- RN 6 east of Ampampamena and south of Ambatomainty.1
- RN 6, north of the turnoff to Belinta.1
- Sadjoavato, forêt de Sahafary. Forêt sèche.1