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- Mantadia National Park, along road going North into the park.2
- Road from Route Nationale 2 to Lakato.2
- Alaotra Mangoro, Antampimbato, Analamay. Forest without cuirasse. Collected with F. Edmond, B.A. Ratodimanana, S. Randrianasolo & J.F. Randriamitantsoa.1
- Alaotra Mangoro, Moramanga, Ambohibary, Ampitambe, Ambatovy. Andranovery edge of forest. Collected with S. Randrianasolo, R. Bernard, B.A. Ratodimanana & F. Edmond.1
- Alaotra Mangoro, Moramanga, Andasibe, Menalamba, Antsahalava. Antsahalava forest. Collected with H.R. Radona, B.A. Ratodimanana, J.F. Randriamitantsoa & A. Rakotondrafara.1
- Ambatondrazaka, Commune Didy, Fokontany Antsevabe, forêt de Sahananto, piste vers Bemainty. Forêt dense humide de moyenne altitude sur pente. Echantillons préservés en alcool. Récolté avec M.F. Randriatsivery, N.M. Andrianjafy & V. Raparison.1
- Ambatovy base camp1
- Ambatovy-Sahavarina Forest1
- Andasibe (Perinet) Forest N.E. of Graphite Mine.1
- Andasibe National Park, in picnic area near visitor center.1
- Andasibe National Park, on trail to the East of visitor center.1
- Mantadia National Park, along trail starting from the northernmost parking area of road going North into park.1
- Moramanga, Andasibe, Analamazaotra-Périnet RS, Circuit Indri 1.1
- Moramanga, Andasibe, Berano. Ambatovy, Ampangadiantrandraka forest.1
- Moramanga, Andasibe, Parc National Analamazaotra-Périnet, Andranomadio.1
- Moramanga, Andasibe, Parc National Mantadia, Falierana, Sahandambo.1
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