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- Firondronana de Mananara-Nord, Ivontaka-Sud; forêt à 100 m de la mer; parcelle 4.1
- Masoala National Park. Ridge NNE of summit of Ambohitsitondroinan'Mahalevona, SSE of village of Mahalevona. Low perhumid forest along trail.1
- Station forestière de Tampolo, 10 km north of Fenoarivo, east of RN 5. Plant preserved in alcohol, collected under MBG second Field Botanist Training program with students from ESSA; part of "Inventaire Biologique Tampolo 1997."1
- Toamasina. Masoala National Park. 1 km NE (straight line) of Andranobe (Park Station).1
